Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White House.

Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Gramma the tortoise, the oldest living resident at the San Diego Zoo, died at the age of 141, according to the zoo.

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved ...

The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died.

The beloved Galapagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has died LOS ANGELES -- After more than a century of munching on her favorite ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Tortoises can live for more than 100 years in the wild and close to double that in captivity. This tortoise was born in her native habitat likely in the 1880s.

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.